B. W.  Price, Villet, M. H., Walton, S. M., and Barker, N. P., ?Using molecules and morphology to infer the phylogenetic relationships and evolutionary history of the Dirini (Nymphalidae: Satyrinae), a tribe of butterflies endemic to Southern Africa?, Systematic Entomology, vol. 36, pp. 300 - 316, 2011.